Children (9 direct children, 9 direct and indirect children):
Handleyomys fuscatus
Dusky-footed handley’s mouse
Handleyomys melanotis
Black-eared oryzomys
Handleyomys saturatior
Cloud forest oryzomys
Handleyomys chapmani
Chapman’s rice rat
Handleyomys intectus
White-footed handley’s mouse
Handleyomys rostratus
Long-nosed oryzomys
Handleyomys rhabdops
Highland oryzomys
Handleyomys alfaroi
Alfaro’s rice rat
Handleyomys guerrerensis