Children (537 direct children, 600 direct and indirect children):
Harpalus tardus
Harpalus affinis
Harpalus rubripes
(tree size 1)
Harpalus caliginosus
Murky ground beetle
Harpalus antonowi
Harpalus semenowi
Harpalus setiporus
Harpalus cephalotes
Harpalus latus
(tree size 1)
Harpalus laevipes
Harpalus flavescens
Harpalus anxius
Harpalus chalcentus
Harpalus honestus
(tree size 1)
Harpalus hospes
(tree size 1)
Harpalus luteicornis
Harpalus modestus
Harpalus pennsylvanicus
Harpalus punctipennis
Harpalus saxicola
Harpalus serripes
(tree size 1)
Harpalus servus
Harpalus zabroides
Harpalus rufipes
Harpalus autumnalis
Harpalus actiosus
Harpalus acupalpoides
Harpalus adenticulatus
Harpalus aeneipennis
Harpalus aenigma
Harpalus aequicollis
Harpalus aesculanus
Harpalus agakhaniantzi
Harpalus akinini
Harpalus alajensis
Harpalus albanicus
Harpalus alexandrae
Harpalus alexeevi
Harpalus alienus
Harpalus alpivagus
Harpalus amarellus
Harpalus amariformis
Harpalus amplicollis
Harpalus amputatus
Harpalus anatolicus
Harpalus angulatus
(tree size 1)
Harpalus angustipennis
Harpalus angustitarsis
Harpalus animosus
Harpalus anisodactyliformis
Harpalus anxioides
Harpalus apache
Harpalus araraticus
Harpalus arcuatus
Harpalus arnoldii
Harpalus ascetes
Harpalus asemus
Harpalus asphaltinus
Harpalus aterrimus
Harpalus atratus
Harpalus atrichatus
Harpalus atripes
Harpalus attenuatus
Harpalus australasiae
Harpalus azumai
Harpalus babai
Harpalus badakschanus
Harpalus baleensis
Harpalus balli
Harpalus basanicus
Harpalus basilewskyi
Harpalus basuto
Harpalus bellieri
Harpalus beneshi
Harpalus bitinctus
Harpalus brachypterus
Harpalus brevicornis
Harpalus brevis
Harpalus breviusculus
Harpalus brunnipes
Harpalus bucharicus
Harpalus bungii
Harpalus caeruleatus
Harpalus caiphus
Harpalus calathoides
Harpalus calceatus
Harpalus capicola
Harpalus cardoni
Harpalus caspius
Harpalus cautus
Harpalus chalcopterus
Harpalus chasanensis
Harpalus chengjiangensis
Harpalus chiloschizontus
Harpalus chobautianus
Harpalus chrysopus
Harpalus cilihumerus
Harpalus circumpunctatus
Harpalus cisteloides
(tree size 2)
Harpalus cohni
Harpalus colchicus
Harpalus compar
Harpalus compressus
Harpalus contemptus
Harpalus convexus
Harpalus cordatus
Harpalus cordifer
Harpalus coreanus
Harpalus corporosus
Harpalus corpulentus
Harpalus corrugatus
Harpalus crates
Harpalus cupreus
(tree size 2)
Harpalus cursorius
Harpalus cyanopterus
Harpalus cyclogonus
(tree size 1)
Harpalus cyrtonotoides
Harpalus danieli
Harpalus davidi
Harpalus davidianus
Harpalus decipiens
Harpalus defector
Harpalus demelti
Harpalus desertus
Harpalus diligens
Harpalus dimidiatus
Harpalus disaogashimensis
Harpalus disimuciulus
Harpalus dispar
(tree size 2)
Harpalus dissitus
Harpalus distinguendus
(tree size 2)
Harpalus diversicollis
Harpalus dubius
Harpalus dudkoi
Harpalus durangoensis
Harpalus ebeninus
Harpalus eberlovi
Harpalus egenus
Harpalus egorovi
Harpalus ellipsis
Harpalus eous
Harpalus eraticus
Harpalus erosus
Harpalus erythropus
Harpalus euchlorus
Harpalus exiguus
Harpalus famelicus
Harpalus farkaci
Harpalus faunus
Harpalus ferghanensis
Harpalus fimetarius
Harpalus flavicornis
(tree size 1)
Harpalus fokienensis
Harpalus foveiger
Harpalus franzi
Harpalus frater
Harpalus fraternus
Harpalus froelichii
Ground beetle
Harpalus fukiensis
Harpalus fulgens
Harpalus fulvicornis
Harpalus fulvilabris
Harpalus fulvipennis
Harpalus fulvus
Harpalus fuscicornis
Harpalus fuscipalpis
Harpalus fuscipennis
Harpalus fuscoaeneus
Harpalus ganssuensis
Harpalus giacomazzoi
Harpalus gilgil
Harpalus gisellae
Harpalus glasunovi
Harpalus gravis
Harpalus gregoryi
Harpalus grilli
Harpalus griseus
Harpalus hartmanni
Harpalus hatchi
Harpalus hauserianus
Harpalus herbivagus
Harpalus heyrovskyi
Harpalus hiekei
Harpalus hirtipes
Harpalus huachuca
Harpalus hybridus
Harpalus hypogeomysis
Harpalus idiotus
Harpalus impressicollis
Harpalus impressus
Harpalus inconcinnus
Harpalus indianus
Harpalus indicola
Harpalus indicus
Harpalus indigens
Harpalus inexspectatus
Harpalus ingenuus
Harpalus innocuus
Harpalus italus
Harpalus janinae
Harpalus jeanneli
Harpalus jordanus
Harpalus jureceki
Harpalus kabakianus
Harpalus kabakovi
Harpalus kadleci
Harpalus kadyrbekovi
Harpalus kagyzmanicus
Harpalus kailiensis
Harpalus kandaharensis
Harpalus karamani
(tree size 2)
Harpalus karokorum
Harpalus katiae
Harpalus kazanensis
Harpalus kaznakovi
Harpalus kibonoti
Harpalus kirgisicus
Harpalus kiritshenkoi
Harpalus klapperichi
Harpalus kmecoi
Harpalus koshiensis
Harpalus kozlovi
Harpalus kryzhanovskii
Harpalus kunarensis
Harpalus laetus
Harpalus lama
Harpalus lampronotus
Harpalus lateralis
Harpalus laticeps
Harpalus lederi
Harpalus lethierryi
(tree size 1)
Harpalus lewisii
Harpalus liobasis
Harpalus litigiosus
Harpalus longicollis
Harpalus longihornus
Harpalus longipalmatus
Harpalus lopezi
Harpalus lugubris
Harpalus lumbaris
Harpalus lutshniki
Harpalus macronotus
Harpalus madagascariensis
Harpalus mairei
Harpalus major
Harpalus makhekensis
Harpalus manas
Harpalus marginellus
Harpalus mariae
Harpalus martini
Harpalus masoreoides
Harpalus massarti
Harpalus mauritanicus
Harpalus medvedevi
Harpalus megacephalus
Harpalus meghalayensis
Harpalus melancholicus
Harpalus melaneus
Harpalus meridianus
Harpalus metallinus
Harpalus metarsius
Harpalus meteorus
Harpalus michaili
Harpalus michailovi
Harpalus microdemas
Harpalus microthorax
Harpalus miles
Harpalus minutulus
Harpalus mitridati
Harpalus mlynari
Harpalus morvani
Harpalus muciulus
Harpalus nanniscus
Harpalus natalensis
Harpalus natalicus
Harpalus neglectus
(tree size 2)
Harpalus nevadensis
Harpalus nigrans
Harpalus nigritarsis
Harpalus numidicus
Harpalus nyassicus
Harpalus obliquus
Harpalus oblitus
(tree size 2)
Harpalus obnixus
Harpalus obtusiusculus
Harpalus ochropus
Harpalus oodioides
Harpalus opacipennis
Harpalus optabilis
Harpalus ovtshinnikovi
Harpalus pachys
Harpalus paivanus
Harpalus pallidipennis
Harpalus pallipes
Harpalus parasinuatus
Harpalus paratus
Harpalus parvulus
Harpalus pastor
Harpalus penglainus
Harpalus pensylvanicus
Pennsylvania dingy ground beetle
Harpalus periglabellus
Harpalus perlucens
Harpalus petri
Harpalus picipennis
Harpalus pilosus
Harpalus plancyi
Harpalus platynotus
Harpalus plenalis
Harpalus politus
(tree size 1)
Harpalus polyglyptus
Harpalus poncei
Harpalus potanini
Harpalus poussereaui
Harpalus praecurrens
Harpalus praticola
Harpalus procognatus
Harpalus progrediens
Harpalus prosperus
Harpalus protractus
Harpalus providens
Harpalus pseudohauserianus
Harpalus pseudophonoides
Harpalus pseudoserripes
Harpalus pseudotinctulus
Harpalus pterostichus
Harpalus puetzi
Harpalus pulchirinulus
Harpalus pulvinatus
(tree size 1)
Harpalus pumilus
Harpalus punctatostriatus
Harpalus puncticeps
Harpalus pusillus
Harpalus pygmaeus
Harpalus raphaili
Harpalus reflexus
Harpalus regressus
Harpalus remboides
Harpalus retractus
Harpalus reversus
Harpalus rivalsi
Harpalus roninus
Harpalus rotundus
Harpalus rougemonti
Harpalus rubefactus
Harpalus rufipalpis
(tree size 5)
Harpalus rufiscapus
Harpalus rufitarsoides
(tree size 1)
Harpalus rufomarginatus
Harpalus rumelicus
Harpalus salinulus
Harpalus salinus
(tree size 3)
Harpalus sanctaehelenae
Harpalus scabripectus
Harpalus schaumii
Harpalus sericatus
Harpalus seyrigi
Harpalus siculus
Harpalus signaticornis
Harpalus silipes
Harpalus simplicidens
Harpalus singularis
Harpalus sinicus
Harpalus sinuatipennis
Harpalus sinuatus
Harpalus smaragdinus
Harpalus smyrnensis
(tree size 1)
Harpalus solitaris
Harpalus somereni
Harpalus somnulentus
Harpalus spadiceus
Harpalus spretus
Harpalus spurius
Harpalus stephani
Harpalus stevenii
Harpalus stevensi
Harpalus stoetznerianus
Harpalus strenuus
Harpalus stricticollis
Harpalus subaeneus
Harpalus subcylindricus
Harpalus subphaedrus
Harpalus subtruncatus
Harpalus suensoni
Harpalus sulphuripes
(tree size 3)
Harpalus sundaicus
Harpalus sushenicus
Harpalus suturangulus
Harpalus szalliesi
Harpalus taciturnus
Harpalus tadorcus
Harpalus tangutorum
Harpalus tarsalis
Harpalus tenebrosus
(tree size 1)
Harpalus terrestris
Harpalus texanus
Harpalus tianschanicus
Harpalus tibeticus
Harpalus tichonis
Harpalus tinctulus
Harpalus tiridates
Harpalus tithonus
Harpalus torosensis
Harpalus torridoides
Harpalus trichophorus
Harpalus tridens
Harpalus triseriatus
(tree size 2)
Harpalus turcicus
Harpalus turmalinus
Harpalus udege
Harpalus uniformis
Harpalus ussuricus
Harpalus ussuriensis
Harpalus vanemdeni
Harpalus venator
Harpalus ventralis
Harpalus vereschaginae
Harpalus vernalis
Harpalus vernicosus
Harpalus viridanus
Harpalus viridicupreus
Harpalus vittatus
Harpalus wadiensis
Harpalus wohlberedti
Harpalus xanthopus
(tree size 1)
Harpalus xinjiangensis
Harpalus yinchuanensis
Harpalus aogashimensis
Harpalus capito
Harpalus imerinae
Harpalus quadratus
Harpalus zhdankoi
Harpalus illeticus
Harpalus clientus
Harpalus probatus
Harpalus nitescens
Harpalus furtivus
Harpalus clandestinus
Harpalus maxwelli
Harpalus curticornis
Harpalus socors
Harpalus vespertinus
Harpalus antiphon
Harpalus fractus
Harpalus mobilis
Harpalus coloradensis
Harpalus provensis
Harpalus alaskensis
Harpalus paululus
Harpalus macilentus
Harpalus fuliginosus
Harpalus seclusus
Harpalus fanaticus
Harpalus mansuetus
Harpalus fallax
Harpalus pleuriticus
Harpalus peritus
Harpalus uteanus
Harpalus celox
Harpalus stupidus
Harpalus futilis
Harpalus intactus
Harpalus pellax
Harpalus carbonatus
Harpalus carolinae
Harpalus funerarius
Harpalus hoppingi
Harpalus pimalicus
Harpalus egregius
Harpalus viduus
Harpalus lewisi
Harpalus lecontei
Harpalus retratus
Harpalus erraticus
Harpalus bicolor
Harpalus rufibarbis
Harpalus albionicus
Harpalus depressicollis
Harpalus stepheni
Harpalus huachua
Harpalus cuncticeps
Harpalus nugax
Harpalus malacus
Harpalus
(tree size 21)
Pseudoophonus
Opadius
Loboharpalus
Baeticoharpalus
Plectralidus
Glanodes
Cephalophonus
Zangoharpalus
Harpalus abasinus
Acardystus
Artabas
Harpalophonus
Licinoderus
Harpalus vagans
Harpalus agnatus
Harpalus cyrenaicus
Harpalus karakorum
Harpalus pecinai
Harpalus pulchrinulus
Harpalus reitteri
Harpalus rhemboides
Harpalus semipunctatus
Harpalus tenuestriatus
Harpalus tjanschanicus
Harpalus abolitus
Harpalus anactus
Harpalus bruckmanni
Harpalus burmeisteri
Harpalus conditus
Harpalus constrictus
Harpalus deletus
Harpalus dilatatus
Harpalus ewaldi
Harpalus excavatus
Harpalus heeri
Harpalus knorri
Harpalus lheritieri
Harpalus liasinus
Harpalus maceratus
Harpalus nuperus
Harpalus obtusus
Harpalus oustaleti
Harpalus pleistocenicus
Harpalus redivivus
Harpalus schlotheimi
Harpalus sinis
Harpalus stierlini
Harpalus stygius
Harpalus tabidus
Harpalus tardigradus
Harpalus ulomaeformis
Harpalus veterum
Harpalus whitfieldi
Harpalus agilis