Popillia
(tree size 320)
Cyphochilus
(tree size 53)
Eucanthidium
Goliathus
(tree size 11)
Goliath beetles
Onitis
(tree size 178)
Allidiostomatinae
Ataenius
(tree size 313)
Dynastinae
(tree size 1794)
Rhinoceros beetles
Cetoniinae
(tree size 2239)
Fruit and flower chafers
Melolonthinae
(tree size 10184)
May beetles and junebugs
Rutelinae
(tree size 964)
Shining leaf chafer
Bubas
(tree size 3)
Ateuchus
(tree size 136)
Dynamopodinae
(tree size 2)
Scybalocanthon
(tree size 29)
Euchirinae
Aclopinae
(tree size 55)
Oniticellus
(tree size 21)
Anomala
(tree size 1184)
Scarabaeinae
(tree size 5749)
Dung beetles
Aphodiinae
(tree size 916)
Aphodiine dung beetles
Christmas beetle
(tree size 41)
Anoplognathus
Euoniticellus
(tree size 19)
Oxysternon
(tree size 12)
Valginae
(tree size 27)
Sericinae
(tree size 187)
Eupleurus
(tree size 3)
Eurysternus
(tree size 55)
Orphninae
(tree size 6)
Pachypodinae
Phaenomeridinae
Tomogonus
(tree size 5)
Acanthonitis
(tree size 5)
Actinophorus
Aeschrotes
Afrocanthon
Afroharoldius
Agamopus
(tree size 5)
Aganhyboma
Allogymnopleurus
(tree size 20)
Allonitis
(tree size 1)
Altonthophagus
Amaecylius
Amietina
(tree size 4)
Amphiceratodon
(tree size 4)
Amphionthophagus
Amphistomus
(tree size 25)
Anisocanthon
(tree size 4)
Anomiopsoides
(tree size 4)
Anomiopsis
Anonthobium
(tree size 6)
Anonychonitis
(tree size 1)
Anoplodrepanus
(tree size 2)
Aphengoecus
(tree size 2)
Aptenocanthon
(tree size 8)
Aptychonitis
(tree size 2)
Arrowianella
Atrichius
Aulacium
Aulacopris
(tree size 3)
Baloghonthobium
(tree size 1)
Bdelyopsis
Bdelyropsis
(tree size 3)
Bohepilissus
(tree size 2)
Bolbites
(tree size 1)
Boletoscapter
(tree size 2)
Boreocanthon
(tree size 9)
Boucomontius
Byrrhidium
(tree size 3)
Caccocnemus
Caccophilus
Caeconthobium
(tree size 1)
Caelontherus
Calhyboma
Cambefortius
(tree size 8)
Cambefortantus
(tree size 8)
Canthochilum
(tree size 23)
Canthonella
(tree size 19)
Canthodimorpha
(tree size 1)
Canthonidia
(tree size 2)
Canthonosoma
(tree size 3)
Canthotrypes
(tree size 1)
Catharsiocopris
(tree size 1)
Cephalodesmius
(tree size 3)
Colobonthophagus
Copridaspidus
(tree size 1)
Coprobius
Coprocanthon
Coprophanaeoides
Coptodactyla
(tree size 17)
Coptorrhina
Cyptochirus
(tree size 4)
Deltepilissus
(tree size 2)
Deltohyboma
(tree size 1)
Deronitis
Diaglyptus
Diasomus
Digitonthophagus
(tree size 2)
Diorygopyx
(tree size 8)
Disphysema
(tree size 1)
Dorbignyolus
(tree size 1)
Drepanocerus
(tree size 4)
Drepanoplatynus
(tree size 1)
Drepanopodus
(tree size 2)
Elassocanthon
Endrodius
Endroedyantus
Enicotarsus
Ennearabdus
(tree size 1)
Epactoides
(tree size 37)
Ephirinus
Epilissus
(tree size 25)
Epionitis
Epirhinus
Eremonthophagus
Eucranium
(tree size 6)
Eudinopus
(tree size 1)
Euhyboma
Eurypodea
Eurysternodes
Eutrichillum
(tree size 3)
Falsignambia
(tree size 1)
Formicdubius
Francmonrosia
Frankenbergerius
(tree size 7)
Freyus
Furconthophagus
Gibbonthophagus
Gilletellus
(tree size 2)
Glaphyocanthon
Glaphyrocanthon
Goniocanthon
Hammondantus
(tree size 1)
Monoplistes
(tree size 6)
Neateuchus
Neocanthidium
Neonitis
(tree size 2)
Neopachysoma
Neosaproecius
(tree size 2)
Nesosisyphus
(tree size 4)
Nesovinsonia
(tree size 1)
Notiophanaeus
Nudipleurus
Odontoloma
(tree size 20)
Oficanthon
(tree size 1)
Onthobium
(tree size 14)
Onthocharis
Onthoecus
Onthohpagus
Onthophagiellus
Onthophahus
Plaesiorrhina
(tree size 4)
Pseudocotalpa
(tree size 4)
Canthon
(tree size 192)
Tumblebugs
Euonthophagus
(tree size 26)
Glyphoderus
(tree size 3)
Nesocanthon
Notopedaria
Aegialiinae
(tree size 80)
Chironinae
(tree size 1)
Eremazinae
(tree size 7)
Jewel scarab
(tree size 115)
Chrysina
Eupoecila
(tree size 7)
Phyllotocus
(tree size 32)
Pseudonthobium
(tree size 9)
Phanaeus
(tree size 82)
Rainbow scarabs
Trichillidium
(tree size 4)
Cleptocaccobius
(tree size 24)
Tesserodoniella
(tree size 2)
Tesserodon
(tree size 19)
Lepanus
(tree size 30)
Deltorhinum
(tree size 7)
Pleuronitis
(tree size 4)
Lophodonitis
(tree size 2)
Saprosites
(tree size 144)
Bradypodidium
(tree size 3)
Mimela
(tree size 186)
Asthenopholis
(tree size 8)
Xinidium
(tree size 4)
Theotimius
(tree size 9)
Apterepilissus
(tree size 4)
Saphobius
(tree size 15)
Scatrichus
(tree size 3)
Pseudopedaria
(tree size 3)
Peckolus
(tree size 3)
Nebulasilvius
(tree size 2)
Martinezidium
(tree size 4)
Scatonomus
(tree size 10)
Milichus
(tree size 15)
Sceliages
(tree size 7)
Zonocopris
(tree size 2)
Larhodius
(tree size 3)
Oruscatus
(tree size 2)
Homocorpis
(tree size 2)
Macropanelus
(tree size 3)
Cretoscarabaeus
(tree size 2)
Sulcophanaeus
(tree size 15)
Paracanthon
(tree size 4)
Sauvagesinella
(tree size 3)
Mentophilus
(tree size 2)
Temnoplectron
(tree size 19)
Pachylomera
(tree size 2)
Pseudignambia
(tree size 2)
Pseudocanthon
(tree size 10)
Pseudocopris
(tree size 2)
Macroderes
(tree size 17)
Streblopus
(tree size 2)
Sylvicanthon
(tree size 18)
Megathopa
(tree size 4)
Scybalophagus
(tree size 6)
Pseudosaproecius
(tree size 12)
Thyregis
(tree size 4)
Malagoniella
(tree size 13)
Diabroctis
(tree size 3)
Melanocanthon
(tree size 4)
Stiptocnemis
(tree size 2)
Heteronitis
(tree size 5)
Pseudochironitis
(tree size 2)
Labroma
(tree size 3)
Pinacotarsus
(tree size 2)
Stiptotarsus
(tree size 3)
Isocopris
(tree size 3)
Onychothecus
(tree size 4)
Janssensantus
(tree size 2)
Chiron
(tree size 11)
Pycnopanelus
(tree size 3)
Endroedyolus
(tree size 1)
Trichius
(tree size 17)
Bee beetle
Micrelaphinis
Xynedria
Termitotroginae
(tree size 14)
Aulonocneminae
Outenikwanus
(tree size 1)
Paracryptocanthon
(tree size 1)
Paraphacosomoides
(tree size 1)
Paroniticellus
(tree size 1)
Paronitis
(tree size 1)
Parvuhowdenius
(tree size 1)
Pedaridium
(tree size 14)
Penalus
(tree size 1)
Pereiraidium
(tree size 1)
Phaedotrogus
(tree size 1)
Pseuduroxys
(tree size 1)
Pteronyx
(tree size 1)
Saphobiamorpha
(tree size 1)
Scaptocnemis
(tree size 1)
Hellaserica
(tree size 1)
Sinapisoma
(tree size 1)
Sulcodrepanus
(tree size 1)
Tanzanolus
(tree size 1)
Tetraechma
(tree size 1)
Tetramereia
(tree size 1)
Tragiscus
(tree size 1)
Tropidonitis
(tree size 1)
Unidentis
(tree size 1)
Vulcanocanthon
(tree size 1)
Walterantus
(tree size 1)
Xenocanthon
(tree size 1)
Yvescambefortius
(tree size 1)
Aposchiza
(tree size 2)
Batesiana
(tree size 1)
Cubidens
(tree size 2)
Dolerotarsa
(tree size 1)
Etiserica
(tree size 1)
Idaeserica
(tree size 1)
Nunoidium
(tree size 1)
Mascarena
(tree size 1)
Mesoserica
(tree size 2)
Nanorhopaea
(tree size 2)
Nedymoserica
(tree size 1)
Parabyrsopolis
(tree size 2)
Pentanomala
(tree size 2)
Philoserica
(tree size 1)
Stomanomala
(tree size 2)
Scythrodes
(tree size 1)
Attavicinus
(tree size 1)
Circellium
(tree size 1)
Coproecus
(tree size 1)
Cretorabaeus
(tree size 1)
Degallieridium
(tree size 1)
Feeridium
(tree size 1)
Heterosyphus
(tree size 1)
Holocanthon
(tree size 1)
Hypocanthidium
(tree size 1)
Janssensellus
(tree size 1)
Leotrichillum
(tree size 1)
Lobateuchus
(tree size 1)
Lobidion
(tree size 1)
Megatharsis
(tree size 1)
Megathoposoma
(tree size 1)
Shangaia
(tree size 1)
Hemicorythoderus
(tree size 1)
Opistocus
(tree size 1)
Ancistroma
(tree size 1)
Lacris
(tree size 1)
Rhyncoserica
(tree size 1)
Trochaloserica
(tree size 2)
Cephaloserica
(tree size 1)
Aethiartrogus
(tree size 1)
Energetia
(tree size 1)
Paulosawaya
(tree size 1)
Heptophyllla
(tree size 1)
Lecantotrogus
(tree size 1)
Homolosternus
(tree size 1)
Macrotetrus
(tree size 1)
Hercites
(tree size 1)
Homaloplia
(tree size 2)
Neosaprosites
(tree size 1)
Pelidnotopsis
(tree size 1)
Aphodiocopris
(tree size 1)
Parbomonyx
(tree size 1)
Rhyncosymmela
(tree size 1)
Acanthosternum
(tree size 1)
Campytrochalus
(tree size 1)
Pseudoisonychus
(tree size 1)
Lachnosterna
(tree size 20)
Latescutella
Sphaerocanthon
Macrophylla
(tree size 6)
Onthopagus
(tree size 1)
Oogeneius
(tree size 2)
Genyoschiza
(tree size 2)
Brachymys
(tree size 2)
Meretricula
(tree size 1)
Cretoscarabaeinae
Prototroginae
Erirhipis
Anatropis
Euphoriaspis
Erirhipidia
Euphorhipis
Isorhipina
Haplophoria
Rhipiphoria
Goraqua
Parascarabaeus
Madateuchus
Neomnematium
Phacosoma (boucomont 1914)
Phacosoma
Bryantella
Knysna
(tree size 4)
Euphoriopsis
Cetonischema
Plusiotis
(tree size 1)
Oxycetonia
(tree size 1)
Parascatonomus
(tree size 10)
Ocnodus
(tree size 1)
Ateuchetus
(tree size 5)
Trichoserica
(tree size 1)
Trichelasmus
(tree size 1)
Tribostethes
(tree size 1)
Teraserica
(tree size 1)
Octotemna
(tree size 1)
Onochirus
(tree size 1)
Otoclinus
(tree size 1)
Bielliana
(tree size 1)
Byrromorpha
(tree size 1)
Iapurea
(tree size 1)
Iarupoides
(tree size 1)
Nataia
(tree size 1)
Nothophanus
(tree size 1)
Chaetoserica
(tree size 1)
Ciocotalpa
(tree size 1)
Clitocephala
(tree size 1)
Costleya
(tree size 1)
Demodema
(tree size 1)
Diasticus
(tree size 1)
Dynamopus
(tree size 1)
Eotrochalus
(tree size 1)
Ataenioides
(tree size 1)
Aegialatis
(tree size 1)
Afrodiasticus
(tree size 1)
Lithoscarabaeinae
Phaenomeris
Scelocopris
Coprophaginae
Saprosotellus
(tree size 1)
Paragynaecoserica
(tree size 1)
Apocamenta
(tree size 1)
Leucoserica
(tree size 2)
Loeblietus
(tree size 1)
Martinezia
(tree size 1)
Microcamenta
(tree size 1)
Miltotrogus
(tree size 1)
Mimotribostethes
(tree size 1)
Passalaphodius
(tree size 1)
Plusiotus
(tree size 1)
Podoserica
(tree size 1)
Poecilodiscus
(tree size 1)
Polylamina
(tree size 1)
Protoleura
(tree size 1)
Pseudammoecius
(tree size 1)
Pseudochlorota
(tree size 1)
Saprosties
(tree size 1)
Sarothermus
(tree size 1)
Simogonious
(tree size 1)
Strehlia
(tree size 1)
Astaenosiagum
(tree size 1)
Epicometis
(tree size 1)
Accrosus
(tree size 3)
Idioschema
Matthewsius
Monteithocanthon