Children (10 direct children, 264 direct and indirect children):
Typhaeus
(tree size 9)
Geotrupini
(tree size 149)
Lethrini
Enoplotrupini
(tree size 37)
Ceratotrupini
(tree size 56)
Baraudia
(tree size 1)
Cretogeotrupini
Geotrupoides
(tree size 2)
Orrhodomala
Chromogeotrupini