Children (14 direct children, 1402 direct and indirect children):
Speleobamini
(tree size 13)
Brachyglutini
(tree size 394)
Bythinini
(tree size 571)
Tychini
(tree size 116)
Proterini
(tree size 90)
Valdini
Arnylliini
(tree size 1)
Barrosellini
(tree size 1)
Cyathigerini
Goniacerini
(tree size 53)
Imirini
(tree size 5)
Iniocyphini
(tree size 128)
Machadoini
(tree size 2)
Pygoxyini
(tree size 14)