Children (22 direct children, 29 direct and indirect children):
Spiny pugolovka
Benthophilus spinosus
Caspian stellate tadpole-goby
Benthophilus leobergius
Caspian tadpole goby
(tree size 4)
Benthophilus macrocephalus
Stellate tadpole-goby
(tree size 2)
Benthophilus stellatus
Granular pugolovka
Benthophilus granulosus
Benthophilus grimmi
(tree size 1)
Benthophilus casachicus
Black sea tadpole-goby
Benthophilus nudus
Benthophilus ctenolepidus
Don tadpole-goby
Benthophilus durrelli
Small-spine tadpole-goby
Benthophilus mahmudbejovi
Benthophilus leptocephalus
Benthophilus ragimovi
Azov tadpole goby
Benthophilus magistri
Benthophilus svetovidovi
Baer pugolovka
Benthophilus baeri
Benthophilus kessleri
Short-snout pugolovka
Benthophilus leptorhynchus
Abdurahmanov's pugolovka
Benthophilus abdurahmanovi
Pinchuk's pugolovka
Benthophilus pinchuki
Benthophilus leobergi
Benthophilus monstrosus