Children (52 direct children, 5749 direct and indirect children):
Ateuchini
(tree size 260)
Canthonini
(tree size 352)
Deltochilini
Coprini
(tree size 917)
Eucraniini
Eurysternini
Oniticellini
(tree size 147)
Onitini
(tree size 39)
Onthophagini
(tree size 2938)
Phanaeini
(tree size 59)
Scarabaeini
(tree size 324)
Aphengium
(tree size 4)
Caccobiomorphus
(tree size 1)
Chalcocopris
(tree size 2)
Chalconotus
(tree size 9)
Deltochilum
(tree size 111)
Dendropaemon
(tree size 49)
Dicranocara
(tree size 4)
Gromphas
(tree size 7)
Mnematium
(tree size 2)
Mnematidium
(tree size 1)
Namakwanus
(tree size 4)
Parachorius
(tree size 20)
Cassolus
Metacatharsius
(tree size 65)
Panelus
(tree size 36)
Latodrepanus
(tree size 6)
Eodrepanus
(tree size 9)
Paraphytus
(tree size 14)
Paragymnopleurus
(tree size 15)
Holocephalus
(tree size 6)
Sinodrepanus
(tree size 8)
Nitiocellus
(tree size 2)
Afrodrepanus
(tree size 2)
Clypeodrepanus
(tree size 3)
Gymnopleurini
(tree size 95)
Sisyphini
(tree size 109)
Paronthobium
(tree size 22)
Proteroscarabaeus
(tree size 6)
Homocopris
(tree size 3)
Hansreia
(tree size 6)
Ignambia
(tree size 2)
Deltochilini
(tree size 14)
Deltorrhinum
(tree size 1)
Coprinisphaera
Lithanomala
(tree size 3)
Hybosorites
(tree size 1)
Epidrepanus
(tree size 3)
Kurtops
(tree size 3)
Microcopris
(tree size 10)
Parateuchus
(tree size 1)
Pleronyx
(tree size 1)
Scaptodera
(tree size 1)
Prionocephale