Children (41 direct children, 1124 direct and indirect children):
Bolboceratinae
(tree size 389)
Geotrupes
(tree size 42)
Geotrupinae
(tree size 264)
Anoplotrupes
(tree size 4)
Bolbotrypes
(tree size 2)
Bolbocerodema
(tree size 8)
Cnemotrupes
(tree size 17)
Geohowdenius
(tree size 3)
Lethrinae
(tree size 141)
Mycotrupes
(tree size 5)
Ceratotrupes
(tree size 3)
Peltotrupes
(tree size 2)
Haplogeotrupes
(tree size 4)
Megatrupes
(tree size 2)
Epigeotrupes
(tree size 4)
Thorectes
(tree size 46)
Onthotrupes
(tree size 10)
Glyptogeotrupes
(tree size 3)
Frickius
(tree size 2)
Bolboleaus
(tree size 15)
Bolborhombus
(tree size 7)
Bolbogonium
(tree size 14)
Pereirabolbus
(tree size 3)
Halffterobolbus
(tree size 3)
Odonteus
(tree size 15)
Blackbolbus
(tree size 31)
Zefevazia
(tree size 5)
Bolboceroides
(tree size 1)
Taurocerastes
(tree size 1)
Allotrupes
(tree size 1)
Halffterius
(tree size 1)
Lithogeotrupes
(tree size 1)
Parageotrupes
(tree size 1)
Pseudotrypocopris
(tree size 1)
Sericotrupes
(tree size 1)
Sinogeotrupes
(tree size 15)
Bolbapium
(tree size 13)
Zuninoeus
(tree size 1)
Allotrypes
(tree size 1)
Cretogeotrupes
(tree size 1)
Taurocerastinae