Children (8 direct children, 917 direct and indirect children):
Heliocopris
(tree size 57)
Copris
(tree size 291)
Black dung beetles
Canthidium
(tree size 178)
Catharsius
(tree size 119)
Dichotomius
(tree size 175)
Ontherus
(tree size 61)
Synapsis
(tree size 24)
Synapsis bates
Litocopris
(tree size 4)