Children (22 direct children, 2938 direct and indirect children):
Onthophagus
(tree size 2519)
Caccobius
(tree size 110)
Caccobius (beetle)
Alloscelus
(tree size 4)
Anoctus
(tree size 5)
Cassolus
(tree size 8)
Eusaproecius
(tree size 7)
Diastellopalpus
(tree size 4)
Haroldius
(tree size 39)
Heteroclitopus
(tree size 10)
Pinacopodius
(tree size 2)
Stiptopodius
(tree size 11)
Cyobius
(tree size 5)
Phalops
(tree size 38)
Mimonthophagus
(tree size 8)
Sukelus
(tree size 1)
Krikkenius
(tree size 1)
Megaponerophilus
(tree size 1)
Hyalonthophagus
(tree size 9)
Copriodes gistl
Copriodes
Hamonthophagus
(tree size 5)
Morettius
(tree size 2)
Proagoderus
(tree size 127)