Children (12 direct children, 25 direct and indirect children):
Geotrupes vernalis
Trypocopris vernalis
Trypocopris amedei
Trypocopris inermis
Trypocopris alpinus
(tree size 4)
Trypocopris pyrenaeus
(tree size 3)
Trypocopris zaitzevi
Trypocopris fulgidus
Trypocopris vernalis
(tree size 6)
Trypocopris caspius
Trypocopris fausti
Pseudotrypocopris
Trypocopris