Children (12 direct children, 18 direct and indirect children):
Black and caspian sea sprat
Clupeonella cultriventris
Abrau sprat
(tree size 2)
Clupeonella abrau
Anchovy sprat
Clupeonella engrauliformis
Southern caspian sprat
Clupeonella grimmi
Caspian tyulka
(tree size 1)
Clupeonella caspia
Freshwater tyulka
Clupeonella tscharchalensis
Clupeonella muhlisi
Clupeonella curensis
Clupeonella delicatula
(tree size 3)
Clupeonella kessleri
Clupeonella leucocephala
Clupeonella sphaerocephala