Children (18 direct children, 26 direct and indirect children):
Squalidus minor
Squalidus argentatus
Squalidus wolterstorffi
Squalidus iijimae
Squalidus atromaculatus
Squalidus nitens
Squalidus banarescui
Squalidus multimaculatus
Squalidus intermedius
Parasqualidus maii
Squalidus homozonus
Squalidus mantschuricus
Squalidus chankaensis
(tree size 4)
Khanka gudgeon
Squalidus gracilis
(tree size 2)
Squalidus japonicus
(tree size 2)
Squalidus baicalensis
Squalidus coreanus
Squalidus majimae